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* '''Sivak''', J.; Howland, H. & McGill-Harelstad, P. (1987) "Vision of the Humboldt Penguin (Spheniscus humboldti) in Air and Water " ''Proceedings of the Royal Society of London. Series B, Biological Sciences''. '''229'''(1257): 467-472
* '''Slack''', Kerryn E.; Jones, Craig M.; Ando, Tatsuro; Harrison G. L. "Abby"; Fordyce R. Ewan; Arnason, Ulfur & Penny, David (2006): Early Penguin Fossils, plus Mitochondrial Genomes, Calibrate Avian Evolution. ''Molecular Biology and Evolution'' '''23'''(6): 1144-1155. <small>{{DOI|10.1093/molbev/msj124}}</small> [http://mbe.oxfordjournals.org/cgi/reprint/23/6/1144 PDF fulltext] [http://mbe.oxfordjournals.org/cgi/content/full/msj124/DC1 Supplementary Material]
* '''Wever''', E.; Herman, P.; Simmons, J. & Hertzler D (1969) "Hearing in the Blackfooted Penguin, Spheniscus demersus, as Represented by the Cochlear Potentials" ''PNAS'' '''63'''(3): 676-680 [http://www.pnas.org/cgi/reprint/63/3/676] {{Семарх|url=https://web.archive.org/web/20080619232756/http://www.pnas.org/cgi/reprint/63/3/676 |date=2008-06-19 }}
* '''Williams'''; Tony D. (1995). ''The Penguins - Spheniscidae'' . Oxford: Oxford University Press. <small>ISBN 0-19-854667-X</small>
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